Epipedobates machalilla Hyloxalus cevallosi H. breviquartus

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(1) SVL, males, 19.5-22.8 (n = 9); females, 21.9-25.2 (x = 24.0; n = 8); (2) disc on Finger III expanded; (3) Finger I = II; (4) fringe present on Finger II; (5) disc on Toe IV expanded; (6) fringe present on Toe IV; (7) outer tarsal fold present; (8) toe webbing formula I2—2 Colostethus sauli is compared with those species having an immaculate white venter and a complete oblique lateral stripe, and lacking discrete marks on the gular-chest region (C. breviquartus, C. cevallosi, and C. machalilla). It is distinguished from all of them by having basal webbing on the toes.

Description and variation.—The original description (Edwards, 1974) is appropriate, except that Fingers I and II are about equal in length, whereas Edwards reported Finger I longer than Finger II; the oblique tarsal fold extends to the margin of toe and is continuous with a lateral fringe (oblique tarsal fold not extending from the inner metatarsal tubercle according to Edwards, 1974) . The dorsum was described as having distinct black blotches (Edwards, 1974) or irregular, but usually transverse, black markings (Duellman, 1978). Usually the dorsum pattern consists of an interorbital square mark, a "V"-shaped mark on the anterior body, and a transverse mark on the posterior part of the body; also a black dorsolateral stripe extends from the anal region to the scapular region. A narrow middorsal gray line occurs in one specimen. A single specimen from the southernmost locality NE Montalvo, in Locación Petrolera Garza 1, Provincia Pastaza, has slightly less webbing than topotypic material. Tadpoles.—Edwards (1974) described free-swimming tadpoles of Stages 27-36. Duellman (1978) described tadpoles in Stage 36. Calls.—Duellman (1978) described the call as a series of two or three quickly repeated peeps.

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