Hyloxalus cevallosi Colostethus fugax Epipedobates machalilla H. breviquartus

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(1) SVL, males unknown in the Ecuadorian sample; females, 15.4-16.8 (x = 16.3; n = 3); (2) disc on Finger III expanded; (3) length of Finger I = II; (4) fringe absent on Finger II; (5) disc on Toe IV expanded; (6) fringe present on Toe IV; (7) outer tarsal fold absent; (8) toe webbing absent; (9) dorsolateral stripe absent; (10) oblique lateral stripe present, extending to eye; (11) ventrolateral stripe absent; (12) discrete marks absent on gular-chest region; (13) abdomen white; (15) third finger not swollen in males (reported for a Colombian population by Rivero and Serna, 1988:147); (16) testes color unknown (Coloma, 1995).Colostethus breviquartus is compared with those species that have a complete oblique lateral stripe, have rudimentary or no webbing on the toes, and have an immaculate white venter (C. cevallosi, C. fugax, and C. machalilla ). It is distinguished from C. fugax and C. machalilla by males lacking a Finger III swollen. Furthermore, C. breviquartus differs from C. machalilla by a different dorsal color pattern (an "X"-shaped mark in C. machalilla and a uniform dorsum in C. breviquartus) and from C. cevallosi by having fringed toes.

Description and variation.—The original description (Rivero and Serna, 1986) is appropriate and corresponds to Ecuadorian specimens examined, except that the first and second fingers are equal in length (Finger I slightly longer than Finger II in holotype), and the largest Ecuadorian specimen is an adult female having a SVL of 16.8 mm, in contrast to the holotype that has a SVL of 21.2 mm. There is variation in dorsal coloration in Ecuadorian specimens. One specimen has small dark flecks on the dorsum, but the others lack flecks. One has a well-defined black band bordering the oblique lateral stripe in contrast to diffuse bands in the others. Also, one has a well-defined transverse band on the dorsal surface of the shank (Coloma, 1995).

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